All Evolutions in Warhammer Survivors

One of the best parts about Warhammer Survivors is finding new evolutions as you unlock new characters. If you want to unlock every evolution and solve their minigames, then we’ve got the evolution formulas you need to know below so you won’t waste time experimenting. Check out the guide below to jump straight back into the action.

All Evolution Formulas in Warhammer Survivors

Image: GameRant via Auroch Digital

Warhammer 40K Universe

Age of Sigmar Universe

Best Crowd Control Evolutions

Image: GameRant via Auroch Digital

Most players want one thing from the evolutions, and that’s just instantly wiping the area of the hordes of enemies. However, you will need multiple evolutions to make it to the boss fight, and we’ll show the best below:

Mordant Earth

Coming from the combo of Nuln Oil and Hourglass, Mordant Earth is by far one of the best and most accessible crowd control evolutions that not only kills enemies in large quantities, but also slows them down significantly. The fact that every character has direct access to this evolution without a follower makes it pretty accessible.

To make Mordant Earth even more effective, you can use the Tape Measure passive skill to increase the size of every drop to its maximum amount. This way, every drop of the black liquid will encompass a large area, damaging and killing even more enemies.

Bandolier and Gauntlet are some other passives that can be useful along with Mordant Earth. The former will increase the drop rate of the black liquid, while the latter raises the base damage rate.

Side Sponsons

The tank itself is already an amazing weapon to crush minions by driving over them, but the evolution gives it two big machine guns that target the areas above and below the tank.

Side Sponsons is an evolution that doesn’t eliminate its base, meaning the Battle Cannon will still remain operational at full force to target the enemies, though that’s not why I love the tank.

Gauntlet is a must-have passive skill for Side Sponsons, and that’s not about the machine gun damage, but rather the tank’s crushing damage. At the base level, the tank deals nearly 300 damage every time you run over an enemy. With a maxed-out Gauntlet passive, this number will go up by nearly 50%, and that’s a massive jump near the end of the missions before facing the bosses. You will definitely need that to stay alive until you can meet the boss, especially in high-tier stages.

Heavy Flamer

If you want to get the most out of this weapon, you need to equip Bandolier and max out its level. Only after that can you burn down your enemies non-stop.

Heavy Flamer does a great job of opening a path for you to move forward. This weapon covers almost 180 degrees in the direction you’re pointing it. The only problem is that the evolution has a long reload time. So, it won’t be a non-stop weapon. And that’s why you need Bandolier to fix that problem.

Keep in mind that no matter how much you tweak Heavy Flamer, it won’t be as powerful as Promethium Canister. So, for defeating elite or extremis bosses, you still need that pick up.

Aquatic Illusions

If you are playing in the Age of Sigmar universe, Aquatic Illusions is what you should focus on. Aquatic Illusions is like a single drop of Mordant Earth, but you carry it around with you.

So, you slow down enemies, and you damage them as long as they are in the radius of evolution. Maxing out Tape Measure and Bandolier will significantly help with the size of this evolution and its hit rate.

Best Boss Damage Evolutions

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Not all the crowd control evolutions work great with bosses. Whenever you are about to fight an Extremis boss, you want to keep a safe distance.

You should have at least one or two evolutions that deal a significant amount of damage to a single target, or you will have a hard time defeating bosses.

Deathstrike Missile

Deathstrike Missile is pretty good at both crowd control and boss damage, but it has a pretty long reload time, which prevents it from being an ideal option for crowd control without other evolutions.

On the other hand, it’s phenomenal for boss damage. This is the only evolution that uses a precise targeting system and always targets the enemy with the highest HP. So, even if you are surrounded by hordes of enemies with an elite boss among them, Deathstrike Missile will directly hit the elite, though its area of effect will still melt a large number of regular enemies too.

If you want to negate a boss’ ability to heal itself, then Deathstrike Missile is a must-have evolution.

Heavy Bolter

Heavy Bolter is a machine gun with a long range that allows you to target enemies no matter how far they are standing from you. Since you are the one who aims Heavy Bolter, it’s always a solid evolution against bosses.

However, it’s not as powerful as other evolutions, so you can’t rely solely on Heavy Bolter. Using Gauntlet can boost its damage output a little, but that still wouldn’t be enough.

Weapon of the Matriarch

This is the only melee evolution that can actually count as a ranged one too. The evolution is a long burning blade, so long that it barely fits on screen, but it can even get better.

If you max out the Tape Measure passive skill, Weapon of the Matriarch will become so big and long that you will only see its hilt on your screen. So, Tape Measure basically makes this blade a great boss battle evolution.

The problem with Weapon of Matriarch is its reload time. It’s almost like the Heavy Flamer. To make it appear on your screen non-stop, you need to max out the Bandolier passive skill as well. With this, you will have three passive slots occupied for the sake of one evolution, which makes a weaker choice than the recommendations above.

Typhoon Crossbow

Another single-shot evolution like Heavy Bolter, but this one comes with automatic aiming. Typhoon Crossbow doesn’t have the insane fire rate of Heavy Bolter, but it automatically shoots bolts at the nearest target, and that means you never miss a shot on a boss.

Typhoon Crossbow is not a Deathstrike Missile. Its damage output is good, but it cannot handle an Extremis boss single-handedly. So, using it along with other evolutions is strongly recommended.

Released

November 10, 2026

ESRB

Everyone 10+ / Fantasy Violence, Mild Blood

Developer(s)

Auroch Digital

Publisher(s)

Auroch Digital